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Fascinating question! I've found some editions of the Aeneid with these extra lines included, and some (most) without. It seems that they aren't found in any of the oldest manuscripts of the Aeneid (except where one commentator scribbled them in the margin much later). Instead, they're first mentioned by the grammarian Aelius Donatus, who wrote in his Vita ...


This rule applies to Russian, Germanic languages, Latin and ancient Greek as far as I know. So I suspect that it's a feature of the proto-IE language.

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