From this answer and Allen & Greenough §493, I understand that Latin does not have a perfect active participle. But on Wiktionary, I see the following usage note in the entry on the suffix -vus:
Originally forming the perfect active participle, as in alvus (“entity having nourished”), clīvus (“entity having leaned”), gnāvus (“having known”).
It's not clear if this is meant to suggest that Latin had a perfect active participle at one point, or if it only existed in Proto-Indo-European.
What is the history of the perfect active participle in Latin? Did Classical Latin or Old Latin ever have such a thing, or did it die out in Proto-Italic or prior?